With the emergence of epigenetic memory, organisms can fix historical and context- dependent impressive experiences. Evolution from now on learnt to learn. Learning means organisms can avoid reproduction of always the same. This is key to adaptation. Epigenetic regulation emerges as a fine-tuned genome-wide network that can rapidly remodel and reprogram genetic content. Epigenetic switching outcompetes genetic mutations (error replications) during adaptation to changing lifeworld. Epigenetic markings can have both short-term and long-term functional effects such as soma to germline inheritance. This symposium assembles approximately 60 experts from different fields to discuss a new paradigmatic understanding of how Evolution learnt to learn. The small number of participants guarantees the legendary relaxed and inspiring atmosphere for presentation and discussion.